Grazing Impacts on Biodiversity, Carbon Cycle, Water Efficiency, and Animal Welfare: Review

Journal title Economia agro-alimentare
Author/s Giulia Pastorelli, Ilaria Falconi, Maria Assunta D'Oronzio, Raffaella Pergamo
Publishing Year 2026 Issue 2025/3
Language English Pages 24 P. 37-60 File size 0 KB
DOI 10.3280/ecag2025oa20804
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<p>This review examines the role of pasture-based livestock systems in promoting agri-environmental sustainability across four key dimensions: biodiversity, water-use efficiency, carbon cycle, and animal welfare. Despite growing political and scientific interest in extensive grazing, the existing literature remains fragmented and limits a comprehensive understanding of grazing’s multidimensional impacts. To address this gap, we conducted a review of studies published between 2010 and 2025, following the Cochrane Handbook and applying the PICO (Population, Intervention, Comparison, Outcome) framework.<br />The analysis focused exclusively on studies conducted in Europe to explore the state of the art on the topic and analyze future policy implications. The results indicate that moderate and well-managed grazing improves biodiversity, increases water-use efficiency, and promotes soil carbon sequestration, especially through rotation practices and tree integration.<br />Access to pasture improves animal welfare, although outcomes vary depending on infrastructure and environmental conditions.<br />The review emphasizes the need for integrated approaches that combine traditional knowledge, environmental design, and precision tools to maximize the sustainability of grazing systems.</p>

Keywords: water-use;carbon cycle;Animal welfare;Biodiversity grazing systems

  1. Agradi, S., Munga, A., Barbato, O., Palme, R., Tarhan, D., Bilgiç, B., Dokuzeylül, B., Ercan, A. M., Or, M. E., Brecchia, G., Curone, G., Draghi, S., Vigo, D., Marongiu, M. L., González-Cabrera, M., & Menchetti, L. (2023). Goat hair as a bioindicator of environmental contaminants and adrenal activation during vertical transhumance. Frontiers in Veterinary Science, 10. Doi: 10.3389/fvets.2023.1274081.
  2. Asheim, L. J., Thorvaldsen, P., & Rivedal, S. (2020). Policy measures to preserve Norwegian coastal and fjord landscapes in small-scale farming systems. Environmental Science and Policy, 104, 43-51. Doi: 10.1016/j.envsci.2019.10.017.
  3. Ates, S., Isik, S., Keles, G., Aktas, A. H., Louhaichi, M., & Nangia, V. (2013). Evaluation of deficit irrigation for efficient sheep production from permanent sown pastures in a dry continental climate. Agricultural Water Management, 119, 135-143. Doi: 10.1016/j.agwat.2012.12.017.
  4. Aubé, L., Mollaret, E., Mialon, M. M., Mounier, L., Veissier, I., & de Boyer des Roches, A. (2023). Measuring the human-animal relationship in cows by avoidance distance at pasture. Applied Animal Behaviour Science, 265. Doi: 10.1016/j.applanim.2023.105999.
  5. Avondo, M., Secchiari, P., Battaglini, L. M., Bonanno, A., & Pulina, G. (2013). Soil, pasture and animal product quality. Italian Journal of Agronomy, 8(3), 141-148. Doi: 10.4081/ija. 2013.e19.
  6. Barron, L. J. R., Andonegi, A., Gamboa, G., Garmendia, E., García, O., Aldai, N., & Aldezabal, A. (2021). Sustainability assessment of pasture-based dairy sheep systems: A multidisciplinary and multiscale approach. Sustainability (Switzerland), 13(7). Doi: 10.3390/su13073994.
  7. Barsotti, M. P., de Almeida, R. G., Macedo, M. C. M., Laura, V. A., Alves, F. v., Werner, J., & Dickhoefer, U. (2022). Assessing the freshwater fluxes related to beef cattle production: A comparison of integrated crop-livestock systems and a conventional grazing system. Agricultural Water Management, 269. Doi: 10.1016/j.agwat.2022.107665.
  8. Battini, M., Renna, M., Giammarino, M., Battaglini, L., & Mattiello, S. (2021). Feasibility and Reliability of the AWIN Welfare Assessment Protocol for Dairy Goats in Semi-extensive Farming Conditions. Frontiers in Veterinary Science, 8. Doi: 10.3389/fvets.2021.731927.
  9. Bernués, A., Olaizola, R. A. c, Villalba, D., & Casasús I. (2011). Sustainability of pasture-based livestock farming systems in the European Mediterranean context: Synergies and trade-offs. Livestock Science, 139(1-2), July, 44-57. Doi: 10.1016/j.livsci.2011.03.018.
  10. Bernués, A., Rodríguez-Ortega, T., Ripoll-Bosch, R., & Alfnes, F. (2014). Sociocultural and economic valuation of ecosystem services provided by Mediterranean mountain agroecosystems. PLoS ONE, 9(7). Doi: 10.1371/journal.pone.0102479.
  11. Beye, H., Taube, F., Donath, T. W., Schulz, J., Hasler, M., & Diekötter, T. (2023). Species Enriched Grass-Clover Pastures Show Distinct Carabid Assemblages and Enhance Endangered Species of Carabid Beetles (Coleoptera: Carabidae) Compared to Continuous Maize. Land, 12(4). Doi: 10.3390/land12040736.
  12. Bosona, T., & Gebresenbet, G. (2021). Multipurpose simulation model for pasturebased mobile Automated Milking and Marketing System, Part-I: Pasture, milk yield, and milk marketing characteristics. Computers and Electronics in Agriculture, 190. Doi: 10.1016/j.compag.2021.106212.
  13. Broom, D. M. (2017). Components of sustainable animal production and the use of silvopastoral systems. Revista Brasileira de Zootecnia, 46(8), 683-688. Doi: 10.1590/S1806-92902017000800009.
  14. Broom, D. M. (2021). A method for assessing sustainability, with beef production as an example. Biological Reviews, 96(5), 1836-1853. Doi: 10.1111/brv.12726.
  15. Broom, D. M. (2022). Animal board invited opinion paper: The use of sustainability scoring to evaluate food production and prepare for the future. Animal, 16(12). Doi: 10.1016/j.animal.2022.100680.
  16. Broom, D. M., Galindo, F. A., & Murgueitio, E. (2013). Sustainable, efficient livestock production with high biodiversity and good welfare for animals. Proceedings of the Royal Society B: Biological Sciences, 280(1771). Doi: 10.1098/rspb.2013.2025.
  17. Candel-Pérez, D., Lucas-Borja, M. E., Plaza-Álvarez, P. A., Carmona Yáñez, M. D., Soria, R., Ortega, R., Miralles, I., Miralha, L., & Zema, D. A. (2024). Effects of grazing on soil properties in mediterranean forests (Central-Eastern Spain). Journal of Environmental Management, 354. Doi: 10.1016/j.jenvman.2024.120316.
  18. Candel-Pérez, D., Lucas-Borja, M. E., Plaza-Álvarez, P. A., Carmona Yáñez, M. D., Soria, R., Ortega, R., Miralles, I., Miralha, L., & Zema, D. A. (2024). Effects of grazing on soil properties in mediterranean forests (Central-Eastern Spain). Journal of Environmental Management, 354. Doi: 10.1016/j.jenvman.2024.120316.
  19. Carswell, A. M., Gongadze, K., Misselbrook, T. H., & Wu, L. (2019). Impact of transition from permanent pasture to new swards on the nitrogen use efficiency, nitrogen and carbon budgets of beef and sheep production. Agriculture, Ecosystems and Environment, 283. Doi: 10.1016/j.agee.2019.106572.
  20. Chará J., Reyes E., Peri P., Otte J., Arce E., Schneider F. (2019). Silvopastoral Systems and their Contri bution to Improved Resource Use and Sustainable Development Goals: Evidence from Latin America. FAO, CIPAV and Agri Benchmark, Cali, 60 pp. Licence: CC BY-NC-SA 3.0 IGO.
  21. Crossley, R. E., Bokkers, E. A. M., Browne, N., Sugrue, K., Kennedy, E., Engel, B., & Conneely, M. (2022). Risk factors associated with the welfare of grazing dairy cows in spring-calving, hybrid pasture-based systems. Preventive Veterinary Medicine, 204. Doi: 10.1016/j.prevetmed.2022.105640.
  22. Crump, A., Jenkins, K., Bethell, E. J., Ferris, C. P., & Arnott, G. (2019). Pasture access affects behavioral indicators of wellbeing in dairy cows. Animals, 9(11). Doi: 10.3390/ani9110902.
  23. Cumpston, M., Flemyng, E., Thomas, J., Higgins, J. P. T., Deeks, J. J., & Clarke, M. J. (2024). Chapter I: Introduction [last updated August 2023]. In: Higgins, J. P. T., Thomas, J., Chandler, J., Cumpston, M., Li, T., Page, M. J., & Welch V. A. (eds.), Cochrane Handbook for Systematic Reviews of Interventions version 6.5. Cochrane. -- Available from training.cochrane.org/handbook.
  24. Delaby, L., Finn, J. A., Grange, G., & Horan, B. (2020). Pasture-Based Dairy Systems in Temperate Lowlands: Challenges and Opportunities for the Future. In Frontiers in Sustainable Food Systems (Vol. 4). Frontiers Media S.A. Doi: 10.3389/fsufs.2020.543587.
  25. Devincenzi, T., Coppa, M., & Cabiddu, A. (2021). Editorial: Ruminant Grazing Behavior: A Tool to Improve Product Quality and Ecosystem Services. In Frontiers in Veterinary Science (Vol. 8). Frontiers Media S.A. Doi: 10.3389/fvets.2021.744200.
  26. Dibari, C., Argenti, G., Catolfi, F., Moriondo, M., Staglianò, N., & Bindi, M. (2015). Pastoral suitability driven by future climate change along the apennines. Italian Journal of Agronomy, 10(3), 109-116. Doi: 10.4081/ija.2015.659.
  27. Dillon P.G. (2018). The evolution of grassland in the European Union in terms of utilisation, productivity, food security and the importance of adoption of technical innovations in increasing sustainability of pasture-based ruminant production systems Northwest Europe: current practices and main agronomic and environmental. Sustainable meat and milk production from grasslands Volume 23
  28. Grassland Science in Europe.
  29. Dumont, B., Rossignol, N., Decuq, F., Note, P., & Farruggia, A. (2020). How does pasture size alter plant-herbivore interactions among grazing cattle?. Grass and Forage Science, 75(4), 438-446. Doi: 10.1111/gfs.12503.
  30. Elorriaga, Y., Marino, D. J., Carriquiriborde, P., & Alicia, R. (2013). Screening of pharmaceuticals in surface water bodies of the Pampas region of Argentina. Int. J. Environ. Res. Public Health, 6, 330-339. Doi: 10.1504/IJENVH.2013.056974.
  31. Enriquez-Hidalgo, D., Teixeira, D. L., Lewis, E., Buckley, F., Boyle, L., & O’Driscoll, K. (2018). Behavioural responses of pasture based dairy cows to short term management in tie-stalls. Applied Animal Behaviour Science, 198, 19-26. Doi: 10.1016/j.applanim.2017.09.012.
  32. Fleurance, G., Farruggia, A., Lanore, L., & Dumont, B. (2016). How does stocking rate influence horse behaviour, performances and pasture biodiversity in mesophile grasslands?. Agriculture, Ecosystems and Environment, 231, 255-263. Doi: 10.1016/j.agee.2016.06.044.
  33. Fracchiolla, M., Terzi, M., D’Amico, F. S., Tedone, L., & Cazzato, E. (2017). Conservation and pastoral value of former arable lands in the agro-pastoral system of the Alta Murgia national park (Southern Italy). Italian Journal of Agronomy, 12(2). Doi: 10.4081/ija.2017.847.
  34. Fraser, M. D., Vallin, H. E., & Roberts, B. P. (2022). Animal board invited review: Grassland-based livestock farming and biodiversity. Animal, 16(12). Doi: 10.1016/j.animal.2022.100671.
  35. Freschi, P., Musto, M., Paolino, R., & Cosentino, C. (2015). Grazing and biodiversity conservation: Highlights on a natura 2000 network site. In: The Sustainability of Agro-Food and Natural Resource Systems in the Mediterranean Basin (pp. 271-288). Springer International Publishing. Doi: 10.1007/978-3-319-16357-4_18.
  36. Futa, B., Ukalska-Jaruga, A., Tajchman, K., Janiszewski, P., & Pecio, M. (2024). Variability in Nutrient Content and Biochemical Parameters of Soil Under Rotational Pasture Management of Farmed Fallow Deer. Agriculture (Switzerland), 14(11). Doi: 10.3390/agriculture14112011.
  37. Gallé, R., Urák, I., Nikolett, G. S., & Hartel, T. (2017). Sparse trees and shrubs confers a high biodiversity to pastures: Case study on spiders from Transylvania. PLoS ONE, 12(9). Doi: 10.1371/journal.pone.0183465.
  38. Grodkowski, G., Gołębiewski, M., Slósarz, J., Grodkowska, K., Kostusiak, P., Sakowski, T., & Puppel, K. (2023). Organic Milk Production and Dairy Farming Constraints and Prospects under the Laws of the European Union. Animals, 13(9). MDPI. Doi: 10.3390/ani13091457.
  39. Hennessy, D., Delaby, L., van den Pol-van Dasselaar, A., & Shalloo, L. (2020). Increasing grazing in dairy cow milk production systems in Europe. Sustainability (Switzerland), 12(6). Doi: 10.3390/su12062443.
  40. Hesselmann, M., Thorne, S., Vitra, A., Steiner, A. K., Leiber, F., & Dittmann, M. T. (2025). Foraging preferences of dairy cows grazing on contrasted multispecies swards. Veterinary and Animal Science, 28. Doi: 10.1016/j.vas.2025.100439.
  41. Hessle, A., & Danielsson, R. (2024). Cattle population required for favorable conservation status of management-dependent semi-natural grasslands and forests, and associated increase in enteric methane emissions. Journal for Nature Conservation, 78. Doi: 10.1016/j.jnc.2024.126571.
  42. Jechalke, S., Heuer, H., Siemens, J., Amelung, W., and Smalla, K. (2014). Fate and effects of veterinary antibiotics in soil. Trends Microbiol., 22, 536-545. Doi: 10.1016/j.tim.2014.05.005.
  43. Kulik, M., Tajchman, K., Lipiec, A., Bąkowski, M., Ukalska-Jaruga, A., Ceacero, F., Pecio, M., & Steiner-Bogdaszewska, Ż. (2023). The Impact of Rotational Pasture Management for Farm-Bred Fallow Deer (Dama dama) on Fodder Quality in the Context of Animal Welfare. Agronomy, 13(4). Doi: 10.3390/agronomy13041155.
  44. Larraz, V., Barrantes, O., & Reiné, R. (2024). Habitat Selection by Free-Grazing Sheep in a Mountain Pasture. Animals, 14(13). Doi: 10.3390/ani14131871.
  45. Lennartsson, T., & Westin, A. (2024). Can Current Grazing Practices Preserve Biodiversity in Semi-natural Pastures? A Study of the Historical Ecology of Swedish Infield Pastures. Martor, 29, 46-67. Doi: 10.57225/martor.2024.29.04.
  46. Marchegiani, S., Gislon, G., Marino, R., Caroprese, M., Albenzio, M., Pinchak, W. E., Carstens, G. E., Ledda, L., Trombetta, M. F., Sandrucci, A., Pasquini, M., Deligios, P. A., & Ceccobelli, S. (2025). Smart technologies for sustainable pasture-based ruminant systems: A review. Smart Agricultural Technology, 10. Doi: 10.1016/j.atech.2025.100789.
  47. McCormick, I. A., & Stokes, J. E. (2023). Stakeholder Challenges and Opportunities of GPS Shock Collars to Achieve Optimum Welfare in a Conservation or Farm Setting. Animals, 13(19). Doi: 10.3390/ani13193084.
  48. Minea, G., Mititelu-Ionuș, O., Gyasi-Agyei, Y., Ciobotaru, N., & Rodrigo-Comino, J. (2022). Impacts of Grazing by Small Ruminants on Hillslope Hydrological Processes: A Review of European Current Understanding. Water Resources Research2024 58(3). Doi: 10.1029/2021WR030716.
  49. Molle, G., Cannas, A., & Gregorini, P. (2022). A review on the effects of part-time grazing herbaceous pastures on feeding behaviour and intake of cattle, sheep and horses. Livestock Science2024 263. Doi: 10.1016/j.livsci.2022.104982.
  50. Morais, T. G., dos Santos, M. P., Barão, L., Domingos, T., & Teixeira, R. F. M. (2025). Grazing or confining – Decoding Beef’s environmental footprint. Environmental Impact Assessment Review, 112. Doi: 10.1016/j.eiar.2025.107846.
  51. Morais, T. G., Teixeira, R. F. M., Rodrigues, N. R., & Domingos, T. (2018). Carbon footprint of milk from pasture-based dairy farms in Azores, Portugal. Sustainability (Switzerland), 10(10). Doi: 10.3390/su10103658.
  52. Moret-Fernández, D., Arroyo, A. I., Herrero, J., Barrantes, O., Alados, C. L., & Pueyo, Y. (2021). Livestock grazing effect on the hydraulic properties of gypseous soils in a Mediterranean region. Catena, 207. Doi: 10.1016/j.catena.2021.105697.
  53. Moret-Fernández, D., Pueyo, Y., Bueno, C. G., & Alados, C. L. (2011). Hydrophysical responses of gypseous and non-gypseous soils to livestock grazing in a semi-arid region of NE Spain. Agricultural Water Management, 98(12), 1822-1827. Doi: 10.1016/j.agwat.2011.07.001.
  54. Niero, G., Bobbo, T., Callegaro, S., Visentin, G., Pornaro, C., Penasa, M., Cozzi, G., de Marchi, M., & Cassandro, M. (2021). Dairy cows’ health during alpine summer grazing as assessed by milk traits, including differential somatic cell count: A case study from Italy. Animals, 11(4). Doi: 10.3390/ani11040981.
  55. O’Brien, D., Bohan, A., McHugh, N., & Shalloo, L. (2016). A life cycle assessment of the effect of intensification on the environmental impacts and resource use of grass- based sheep farming. Agricultural Systems, 148, 95-104. Doi: 10.1016/j.agsy.2016.07.004.
  56. O’Driscoll, K., Lewis, E., & Kennedy, E. (2019). Effect of feed allowance at pasture on the lying behaviour of dairy cows. Applied Animal Behaviour Science, 213, 40-46. Doi: 10.1016/j.applanim.2019.02.002.
  57. O’Leary, N. W., Byrne, D. T., Garcia, P., Werner, J., Cabedoche, M., & Shalloo, L. (2020). Grazing cow behavior’s association with mild and moderate lameness. Animals, 10(4). Doi: 10.3390/ani10040661.
  58. Opio, C., Gerber, P., Mottet, A., Falculli, A., Tempio, G., MacLeod, M. et al. (2013). Greenhouse gas emissions from ruminant supply chains - A global life cycle assessment. Food and Agriculture Organization of the United Nations (FAO), 214.
  59. Page, M. J., McKenzie, J. E., Bossuyt, P. M., Boutron, I., Hoffmann, T. C., Mulrow, C. D., & Moher, D. (2021). The PRISMA 2020 statement: An updated guideline for reporting systematic reviews. BMJ, 372(71). Doi: 10.1136/bmj.n71.
  60. Pauler, C. M., Homburger, H., Lüscher, A., Scherer-Lorenzen, M., & Schneider, M. K. (2025). Ecosystem services in mountain pastures: A complex network of site conditions, climate and management. Agriculture, Ecosystems and Environment, 377. Doi: 10.1016/j.agee.2024.109272.
  61. Phelps, L. N., & Kaplan, J. O. (2017). Land use for animal production in global change studies: Defining and characterizing a framework. Global Change Biology, 23(11), 4457-4471. Doi: 10.1111/gcb.13732.
  62. Plieninger, T., Hartel, T., Martín-López, B., Beaufoy, G., Bergmeier, E., Kirby, K., Montero, M. J., Moreno, G., Oteros-Rozas, E., & van Uytvanck, J. (2015). Woodpastures of Europe: Geographic coverage, social-ecological values, conservation management, and policy implications. Biological Conservation, 190, 70-79. Doi: 10.1016/j.biocon.2015.05.014.
  63. Ponzetta, M. P., Grohmann, F., & Argenti, G. (2010). Effects of two restoration practices on the vegetation of a semi-natural grassland invaded by bracken in Central Italy. Italian Journal of Agronomy, 5(3), 233-240. Doi: 10.4081/ija.2010.233.
  64. Potenza, C., & Fedele, V. (2011). Effectiveness of the GAEC cross-compliance standard protection of permanent pasture in relation to grazing and pasture conservation management in marginal mountain areas. Italian Journal of Agronomy, 6(Supl.1), 83-86. Doi: 10.4081/ija.2011.6.s1.e11.
  65. Ripamonti, A., Mantino, A., Annecchini, F., Cappucci, A., Casarosa, L., Turini, L., Foggi, G., & Mele, M. (2023). Outcomes of a comparison between pastoral and silvopastoral management on beef cattle productivity, animal welfare and pasture depletion in a Mediterranean extensive farm. Agroforestry Systems, 97(6), 1071-1086. Doi: 10.1007/s10457-023-00848-w.
  66. Ripoll-Bosch, R., de Boer, I. J. M., Bernués, A., & Vellinga, T. v. (2013). Accounting for multi-functionality of sheep farming in the carbon footprint of lamb: A comparison of three contrasting Mediterranean systems. Agricultural Systems, 116, 60-68. Doi: 10.1016/j.agsy.2012.11.002.
  67. Rivera, D., Verde, A., Fajardo Rodríguez, J., Ríos, S., Alcaraz, F., Cárceles, C., Ortíz, J., Valdés, A., Ruíz-Gallardo, J. R., García-Flores, A., Palazón, J. A., & Obón, C. (2022). Ethnoveterinary Medicine and Ethnopharmacology in the Main Transhumance Areas of Castilla-La Mancha (Spain). Frontiers in Veterinary Science, 9. Doi: 10.3389/fvets.2022.866132.
  68. Rodriguez, J. (2024). One Health Ethics and the Ethics of Zoonoses: A Silent Call for Global Action. Veterinary Sciences, 11(9), 394. Doi: 10.3390/vetsci11090394.
  69. Roeder, N., Lederbogen, D., Trautner, J., Bergamini, A., Stofer, S., & Scheidegger, C. (2010). The impact of changing agricultural policies on jointly used rough pastures in the Bavarian Pre-Alps: An economic and ecological scenario approach. Ecological Economics, 69(12), 2435-2447. Doi: 10.1016/j.ecolecon.2010.07.013.
  70. Rutherford, N. H., Lively, F. O., & Arnott, G. (2021). A Review of Beef Production Systems for the Sustainable Use of Surplus Male Dairy-Origin Calves Within the UK. Frontiers in Veterinary Science, 8. Doi: 10.3389/fvets.2021.635497.
  71. Sala, V., Moja, L., Moschetti, I., Bidoli, S., Pistotti, V., & Liberati, A. (n.d.). Revisioni sistematiche-Breve guida all’uso.
  72. Sargeant, J. M., Amezcua, M. D. R., Rajic, A., & Waddell, L. (2005). Guide to conducting systematic reviews in agri-food public health. Food Safety Research and Response Network. Supported by the United States Department of Agriculture, Cooperative Research, Education and Extension Service.
  73. Schmitz, A., & Isselstein, J. (2020). Effect of grazing system on grassland plant species richness and vegetation characteristics: Comparing horse and cattle grazing. Sustainability (Switzerland), 12(8). Doi: 10.3390/SU12083300.
  74. Simionesei, L., Ramos, T. B., Oliveira, A. R., Jongen, M., Darouich, H., Weber, K., Proença, V., Domingos, T., & Neves, R. (2018). Modeling soilwater dynamics and pasture growth in the montado ecosystem using MOHID land. Water (Switzerland), 10(4). Doi: 10.3390/w10040489.
  75. Sitzia, T., Semenzato, P., & Trentanovi, G. (2010). Natural reforestation is changing spatial patterns of rural mountain and hill landscapes: A global overview. In Forest Ecology and Management, 259(8), 1354-1362). Doi: 10.1016/j.foreco.2010.01.048.
  76. Smetanová, A., Follain, S., David, M., Ciampalini, R., Raclot, D., Crabit, A., & le Bissonnais, Y. (2019). Landscaping compromises for land degradation neutrality: The case of soil erosion in a Mediterranean agricultural landscape. Journal of Environmental Management, 235, 282-292. Doi: 10.1016/j.jenvman.2019.01.063.
  77. Steiner, M., Öckinger, E., Karrer, G., Winsa, M., & Jonsell, M. (2016). Restoration of semi-natural grasslands, a success for phytophagous beetles (Curculionidae). Biodiversity and Conservation, 25(14), 3005-3022. Doi: 10.1007/s10531-016-1217-4.
  78. Strand, G. H., Hansen, I., de Boon, A., & Sandström, C. (2019). Carnivore Management Zones and their Impact on Sheep Farming in Norway. Environmental Management, 64(5), 537-552. Doi: 10.1007/s00267-019-01212-4.
  79. Szyszko-Podgórska, K., Szweda, Ż., Świątek, M., Ukalska, J., Pietrasz, K., Pietrasz, M., Wilk, P., Orlińska-Woźniak, P., Szalińska, E., Rokicki, T., Tylkowski, S., & Niżnikowski, R. (2024). Impact of Land Use on Peat Soil Elemental Content and Carabidae and Plant Species Composition and Abundance. Sustainability (Switzerland), 16(11). Doi: 10.3390/su16114420.
  80. Tittonell, P. (2021). Beyond CO2: Multiple ecosystem services from ecologically intensive grazing landscapes of South America. Frontiers in Sustainable Food Systems, 5. Doi: 10.3389/fsufs.2021.664103.
  81. Tölgyesi, C., Kelemen, A., Bátori, Z., Kiss, R., Hábenczyus, A. A., Havadtői, K., Varga, A., Erdős, L., Frei, K., Tóth, B., & Török, P. (2023). Maintaining scattered trees to boost carbon stock in temperate pastures does not compromise overall pasture quality for the livestock. Agriculture, Ecosystems and Environment, 351. Doi: 10.1016/j.agee.2023.108477.
  82. Tonn, B., Densing, E. M., Gabler, J., & Isselstein, J. (2019). Grazing-induced patchiness, not grazing intensity, drives plant diversity in European low-input pastures. Journal of Applied Ecology, 56(7), 1624-1636. Doi: 10.1111/1365-2664.13416.
  83. Toro-Mujica, P. M., Aguilar, C., Vera, R., Barba, C., Rivas, J., & García-Martínez, A. (2015). Changes in the pastoral sheep systems of semi-arid Mediterranean areas: Association with common agricultural policy reform and implications for sustainability. Spanish Journal of Agricultural Research, 13(2), 1-11. Doi: 10.5424/sjar/2015132-6984.
  84. Tsiobani, E. T., Yiakoulaki, M. D., Hasanagas, N. D., & Antoniou, I. E. (2023). Water buffalo social networks during their activities on pasture under the rain. Turkish Journal of Veterinary and Animal Sciences, 47(6), 576-583. Doi: 10.55730/1300-0128.4327.
  85. Turini, L., Foggi, G., Mantino, A., Gasparoni, E., Vichi, F., Silvi, A., Armenia, G., Sala, G., Bonelli, F., Sgorbini, M., & Mele, M. (2025). Changes in hematological and hematochemical parameters in lactating dairy sheep according to different pasture management and heat stress risk: A longitudinal study. Veterinary and Animal Science, 27. Doi: 10.1016/j.vas.2024.100419.
  86. van Klink, R., Nolte, S., Mandema, F. S., Lagendijk, D. D. G., WallisDeVries, M. F., Bakker, J. P., Esselink, P., & Smit, C. (2016). Effects of grazing management on biodiversity across trophic levels – The importance of livestock species and stocking density in salt marshes. Agriculture, Ecosystems and Environment, 235, 329-339. Doi: 10.1016/j.agee.2016.11.001.
  87. Vargas-Canales, J. M., Orozco-Cirilo, S., Estrada, S., del Carpio-Ovando, P. S., Camacho-Vera, J. H., López-Carmona, D., García-Melchor, N., Rodríguez-Haros, B., Valdés-Cobos, A., Sánchez-Torres, Y., Fresnedo-Ramírez, J., Palacios-Rangel, M. I., Ocampo-Ledesma, J. G., Barrera-Perales, O. T., Pineda-Pineda, J., Kreimer, P., García-Cruz, J. C., Reyes-Barrera, D. M., Montiel-Flores, J. C., … de Gortari-Rabiela, R. (2024). Science, technology, agri-food systems, health, and wellbeing: logic, dynamics, and relationships. Frontiers in Sustainable Food Systems, 8. Doi: 10.3389/fsufs.2024.1344357.
  88. Verduna, T., Blanc, S., Merlino, V. M., Cornale, P., & Battaglini, L. M. (2020). Sustainability of Four Dairy Farming Scenarios in an Alpine Environment: The Case Study of Toma di Lanzo Cheese. Frontiers in Veterinary Science, 7. Doi: 10.3389/fvets.2020.569167.
  89. Wróbel, B., Zielewicz, W., & Staniak, M. (2023). Challenges of Pasture Feeding Systems – Opportunities and Constraints. Agriculture (Switzerland), 13(5). Doi: 10.3390/agriculture13050974.
  90. Zhu, S., Olde, L., Lewis, K., Quaife, T., Cardenas, L., Loick, N., Xu, J., & Hill, T. (2023). Eddy covariance fluxes over managed ecosystems extrapolated to field scales at fine spatial resolutions. Agricultural and Forest Meteorology, 342. Doi: 10.1016/j.agrformet.2023.109675.

Giulia Pastorelli, Ilaria Falconi, Maria Assunta D'Oronzio, Raffaella Pergamo, Grazing Impacts on Biodiversity, Carbon Cycle, Water Efficiency, and Animal Welfare: Review in "Economia agro-alimentare" 3/2025, pp 37-60, DOI: 10.3280/ecag2025oa20804